Myriophyllum propinquum

Features

Weak aquatic or semiaquatic or terrestrial herb (50-)100-250(-400) mm tall; stems weak, slender, 1.0-1. 5(-3.0) mm diameter, rooting at lower nodes; glabrous. Leaves distinctly dimorphic. Submerged leaves in whorls of 3-4(-5), ovate to orbicular in outline, (6-)10-20(-25) × 10-16(-25) mm, pectinate with 10-22 filiform pinnae 5-6(-20) mm long, with distinct petiole 1-2 mm long. Emergent leaves in whorls of (2-)3-4(-5), linear, (2-)5-7(-12) × 0.5-1.0(-1.5) mm, usually flattened but occasionally terete to filiform, spreading or slightly curved upwards, margin entire or sometimes weakly toothed. Rarely, all or most emergent leaves pinnatifid, 4-5 ×1.5-3.5 mm wide with pinnae to 1.5 mm long. Transitional leaves usually 1-2(-4) whorls only. Inflorescence an indeterminate spike with the unisexual flowers borne singly in axils of the emergent leaves. Each whorl contains flowers of 1 sex only, male in upper 1-8 whorls, female in lower 2-6 whorls, with usually 1-2 sterile whorls between. Occasionally all whorls (up to 20) contain female flowers only. All flowers subtended by 2 bracteoles. Bracteoles ovate, (0.4-)0.5-0.8 × (0.2-)0.4-0.5 mm wide, slightly laciniate towards tip or entire. Male flowers 4-merous, sessile or becoming shortly pedicellate. Sepals 4, ovate, 0.4-0.6 × 0.3-0.4 mm, weakly toothed near apex. Petals 4, hooded, very weakly keeled, 1.8-2.7(-3.0) × 1.4 mm, becoming reflexed and inrolled after anthesis. Stamens 8; filaments 1-2 mm long, elongating after anthesis; anthers yellow, linear-oblong, 1.7-2.4 × 0.4-0.5 mm, antisepalous anthers c. 0.2 mm longer than antipetalous ones. Styles 0. Ovary vestigial. Female flowers 4-merous, sessile. Sepals, petals and stamens 0. Styles 4, clavate, 0.2-0.3 mm long, stigmas occupying most of length of style, white, fimbriate. Ovary 4-merous, oblong, 0.7-0.8 mm long (excluding styles), 0.7 mm wide; carpels cylindrical, smooth, with styles on outermost corners. Fruit pale yellow-brown, cylindrical, sessile, splitting at maturity into 4 mericarps. Mericarps 0.8-1.1 mm long, 0.4-0.5 mm diameter, smooth apart from scattering of very small asperities on outer face, crowned by persistent base of style.