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  4. Piper excelsum subsp. excelsum

Piper excelsum subsp. excelsum

Hemi Matenga Scenic Reserve, Waikanae.<br>Photographer: Jeremy R. Rolfe, Date taken: 15/02/2010, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Male inflorescence. Silverstream, Upper Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 07/10/2006, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Kawakawa.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Kawakawa.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Fruit.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Kawakawa.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Leaf.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Stokes Valley, Lower Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 24/04/2002, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Wellington Botanic Garden.<br>Photographer: Jeremy R. Rolfe, Date taken: 28/01/1986, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Kawakawa.<br>Photographer: Wayne Bennett, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Ripe fruit. Wanganui.<br>Photographer: Colin C. Ogle, Date taken: 26/01/2010, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Ripe and unripe fruit. Wanganui.<br>Photographer: Colin C. Ogle, Date taken: 26/01/2010, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>.
Male inflorescence. Tamahunga, Omaha.<br>Photographer: Jeremy R. Rolfe, Date taken: 14/10/2009, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Male inflorescence. Tamahunga, Omaha.<br>Photographer: Jeremy R. Rolfe, Date taken: 14/10/2009, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Hemi Matenga Scenic Reserve, Waikanae.<br>Photographer: Jeremy R. Rolfe, Date taken: 15/02/2010, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Hemi Matenga Scenic Reserve, Waikanae.<br>Photographer: Jeremy R. Rolfe, Date taken: 15/02/2010, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Western Hutt hills.<br>Photographer: Jeremy R. Rolfe, Date taken: 28/07/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Cathedral Cove, Coromandel.<br>Photographer: John Sawyer, Date taken: 01/08/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Cathedral Cove, Coromandel.<br>Photographer: John Sawyer, Date taken: 01/08/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Piha, west Auckland.<br>Photographer: John Sawyer, Date taken: 01/07/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Near Raglan, west coast, North Island.<br>Photographer: John Sawyer, Date taken: 01/07/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Near Raglan, west coast, North Island.<br>Photographer: John Sawyer, Date taken: 01/07/2013, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
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Common name

kawakawa, pepper tree

Synonyms

Macropiper excelsum (G.Forst.) Miq. subsp. excelsum

Family

Piperaceae

Authority

Piper excelsum G.Forst.

Flora category

Vascular – Native

Endemic taxon

Yes

Endemic genus

No

Endemic family

No

Structural class

Trees & Shrubs - Dicotyledons

NVS code

The National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.

PIPESE

Chromosome number

2n = 26

Current conservation status

  • Conservation status of New Zealand indigenous vascular plants, 2017

The threat classification status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2017 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website This report includes a statistical summary and brief notes on changes since 2012 and replaces all previous NZTCS lists for vascular plants. Authors: By Peter J. de Lange, Jeremy R. Rolfe, John W. Barkla, Shannel P. Courtney, Paul D. Champion, Leon R. Perrie, Sarah M. Beadel, Kerry A. Ford, Ilse Breitwieser, Ines Schönberger, Rowan Hindmarsh-Walls, Peter B. Heenan and Kate Ladley. Please note, threat classifications are often suggested by authors when publications fall between NZTCS assessment periods – a suggested threat classification status has not been assessed by the NZTCS panel.

Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.

2017 | Not Threatened

Previous conservation statuses

2012 | Not Threatened | Qualifiers: Sp

2009 | Not Threatened

2004 | Not Threatened

Brief description

Fleshy shrub with jointed dark twigs bearing large dark green glossy heart-shaped leaves with hard green flower spikes inhabiting south to Banks Peninsula. Leaves to 120mm wide, veins radiating from middle, peppery to taste, often with insect holes. Fruit orange.

Distribution

Endemic. North and South Islands. Common from te Paki south to about Okarito, North Canterbury and Banks Peninsula.

Habitat

Coastal to lowland (extending up 500 m a.s.l. in warmer parts of the country). Usually an important understorey species in coastal forest.

Wetland plant indicator status rating

Information derived from the revised national wetland plant list prepared to assist councils in delineating and monitoring wetlands (Clarkson et al., 2021 Manaaki Whenua – Landcare Research Contract Report LC3975 for Hawke’s Bay Regional Council). The national plant list categorises plants by the extent to which they are found in wetlands and not ‘drylands’. The indicator status ratings are OBL (obligate wetland), FACW (facultative wetland), FAC (facultative), FACU (facultative upland), and UPL (obligate upland).

UPL: Obligate Upland

Rarely is a hydrophyte, almost always in uplands (non-wetlands).

Features

Small tree to at least 5 m tall; stems erect (occasionally layering), not notably lenticellate, new shoots red-green or green (leaf nerves, petioles and new sterns with reddish colouring), taste peppery; pith of axes (including rachis of spike) without a mucilage core. Prophyll a collar to 0.3 (-2.2) mm high. Leaf blades submembranous, orbicular, suborbicular, at vegetative nodes to 100(-120) mm diameter, usually with 5-8 principal nerves, cordate at base, with a very narrow or closed sinus, occasionally basal lobes overlapping, upper surface of blade not bullate; petiole to 40(-60) mm long, c.0.4×as long as blade, the sheath 0.3-1.0(-2)× as long as non-sheathing part, truncate-rounded at apex and not produced there, the non-sheathing part of petiole to 4.0 mm diameter. Inflorescences solitary or 2-3 together on a short (rarely more than 10 mm long) axillary shoot, and (usually solitary) on the adjacent terminal shoot (occasionally this shoot not fertile); reduced leaf at apex of fertile shoot with a glabrous petiole and usually with a green oblong lamina at least 5 mm long, but lamina often ± lacking, especially on terminal fertile shoot. Female inflorescence erect in flowering and remaining so into fruit, peduncle to c. 1.5 cm long, spike to 60(-100) × c.6 mm diameter, with uniseriate usually 5-10-cellular hairs to 0.15 mm long on lower part of bract stalks and sparingly on rachis, these hairs not obvious on the peduncle just below the lowermost bracts; bracts peltate, bract heads 0.40-0.75 mm diameter; flowers at full emergence centred c.1.3 mm apart, emergent part of ovary ovoid; stigmas 3-4(-5), together c. 1.2 mm diameter. Male inflorescence erect, spike to c.110 mm long, proximally c.6 mm diameter, bracts and hairs as in female inflorescence; staminal filaments c. 0.25 mm long, anthers c.1.00 × 0.75 mm wide. Ripe infructescence c.10 mm diameter; fruitlets coalescent, sunken apically about the persistent dark stigmas, exocarp and mesocarp orange; seed oblong to slightly obovoid, apiculate at apex, c.2.0 × 1.5 dark brown, with (3-)4-5(-7) broad longitudinal furrows.

Similar taxa

Distinguished from the other subspecies by the wine-red to purple-black stems and petioles, and by the leaves which are never peltate, though on some of the Hauraki Gulf Islands, Tuhua (Mayor Island) and Chatham Islands plants often have overlapping leaf bases.

Flowering

August - November

Fruiting

Throughout the year

Propagation technique

Easily grown from semi-hardwood cuttings and fresh seed. An attractive small tree that does best in dappled light, within a free draining but permanently moist soil. Very shade tolerant. Cold sensitive and will not tolerate frost.

Etymology

piper: Pepper

excelsum: Tall

Cultural Use/Importance

Kawakawa is a member of the pepper family related to kava - used as a drink in the Pacific. The fruit and leaves are aromatic and Maori used leaves as poultices to treat bruises, and as a sign of mourning.

Foraging for kawakawa

Click on the Radio New Zealand National logo to listen to This Way Up. Simon Morton interviews Johanna Knox about foraging for kawakawa (duration: 12′13″)

Taxonomic Notes

The generic distinction between Macropiper and Piper has always been tenuous. Recently Jaramillo et al. (2008) have shown that Macropiper should be merged in Piper. However, they did not effect the full transfer of the New Zealand taxa to Piper. This action was taken by de Lange (2012) for Macropiper excelsum subsp. psittacorum, Macropiper excelsum subsp. peltatum f. peltatum and f. delangei.

Attribution

Fact sheet prepared for NZPCN by P.J. de Lange 30 August 2005. Description based on Gardner (1997).

References and further reading

de Lange, P.J. 2012: Taxonomic notes on the New Zealand flora: new names in Piper (Piperaceae). New Zealand Journal of Botany DOI:10.1080/0028825X.2012.708904

Gardner, R.O. 1997: Macropiper (Piperaceae) in the south-west Pacific. New Zealand Journal of Botany 35: 293-307.

Jaramillo, M.A.; Callejas, R; Davidson, C.; Smith, J.F.; Stevens, A.C.; Tepe, E.J. 2008: A phylogeny of the tropical genus Piper using ITS and the chloroplast intron psbJ-petA. Systematic Botany 33: 647-660.

NZPCN Fact Sheet citation

Please cite as: de Lange, P.J. (Year at time of access): Piper excelsum subsp. excelsum Fact Sheet (content continuously updated). New Zealand Plant Conservation Network. https://www.nzpcn.org.nz/flora/species/piper-excelsum-subsp-excelsum/ (Date website was queried)

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