Alectryon excelsus subsp. grandis
Common names
Three Kings tītoki
Biostatus
Native – Endemic taxon
Category
Vascular
Structural class
Trees & Shrubs - Dicotyledons
Simplified description
Small tree with spreading branches and a dark fluted trunk of the Three Kings Islands. Leaves with 2-4 offset pairs of glossy dark green leaflets. Flowers red, in small, clustered, sprays. Fruit fleshy red partly surrounding a black seed and expanding from a furry brown capsule.
Flower colours
Red/Pink
Detailed description
Small spreading shrub tree 2-7 x 3-10 m; multitrunked from base. Branches stout, erect, all parts invested with fine, velutinous, ferrugineous hairs. Bark brown, lenticillate. Adult leaves light to dark green, vernicose, imparipinnate, alternate 14-70 mm long. Leaflets 2-4 pairs; lamina 90-160 x 60-90 mm, subcoriaceous, broadly oblong or ovate, apex obtuse, subacute often acuminate, base cuneate, truncate to oblique, upper leaf surface vernicose, bullate; lamina margin entire. Inflorescences axillary 90-120 mm long, sparingly branched panicles. Flowers bisexual or staminate. Petals absent. Stamens 5-8 in bisexual and 6-10 in staminate flowers, crimson. Stigma ovoid, in staminate flowers ovary tholiform, style absent, in perfect flowers broadly urceolate, style 1.5-2 mm, erect. Fruits sessile, 1-2-lobed, 15-20 x 10-14 mm, pubescent, globular, carina 3-5 mm long on one side. Seed 8-10 x 6-9 mm, subglobose, black, lustrous, sarcotesta fleshy, scarlet, papillose.
Similar taxa
Titoki (Alectryon excelsus Gaertn. subsp. excelsus) could be confused with Three Kings Titoki. However, this subspecies is much taller (up to 15 m tall), and usually has a single trunk. The compound leaves have (3-)5(-7) pairs of smooth edged (or toothed) generally narrower leaflets whose surfaces are matt or glossy but never bullate. There are no major differences in the flowers or the fruits.
Distribution
Endemic to the Three Kings Archipelago, where it is known from Great (Manawa Tawhi) and West Islands.
Habitat
Coastal Forest, where it often grows along the more exposed margins. Occasionally it is found as a subcanopy tree within the dense kanuka (Kunzea aff. ericoides) forest that dominates the Three Kings Islands.
Current conservation status
The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2022-2023 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website. This report includes replaces all previous NZTCS lists for vascular plants. Previous assessments can be found here.
- Conservation status of vascular plants in Aotearoa New Zealand, 2023. 2024. Peter J. de Lange, Jane Gosden, Shannel P. Courtney, Alexander J. Fergus, John W. Barkla, Sarah M. Beadel, Paul D. Champion, Rowan Hindmarsh-Walls, Troy Makan and Pascale Michel Department of Conservation. Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.
2023 | Threatened – Nationally Vulnerable | Qualifiers: CD, DPS, IE, RC
Threats
In the past this tree was virtually eliminated by goats. Since goats were eradicated in 1946, Three Kings titoki has gradually increased in range, and is now found in scattered sites across most of the eastern side of Great Island. A single tree has also been reported from West Island.
Detailed taxonomy
Family
Synonyms
Alectryon excelsus var. grandis Cheeseman, Alectryon grandis (Cheeseman) Cheeseman
Endemic taxon
Yes
Endemic genus
No
Endemic family
No
Ecology
Flowering
October to December
Fruiting
December to April
Propagation technique
Fresh seed germinates readily. However, as most seed comes from trees planted nearby titoki (A. excelsus subsp. excelsus) it is difficult to obtain pure seed, and many plants sold by nurseries are F1 hybrids. Hybrids are now commonly seen in street plantings around Auckland City.
Other information
Etymology
alectryon: In Greek mythology Alectryon was punished and turned into a rooster by Ares after failing to keep watch, possible refers to a cockscomb
excelsus: Tall
grandis: Big
NVS code
The National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.
ALEESG
Chromosome number
2n = 32
Previous conservation statuses
The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2022-2023 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website. This report includes replaces all previous NZTCS lists for vascular plants. Previous assessments can be found here.
- Conservation status of vascular plants in Aotearoa New Zealand, 2023. 2024. Peter J. de Lange, Jane Gosden, Shannel P. Courtney, Alexander J. Fergus, John W. Barkla, Sarah M. Beadel, Paul D. Champion, Rowan Hindmarsh-Walls, Troy Makan and Pascale Michel Department of Conservation. Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.
2017 | Threatened – Nationally Vulnerable | Qualifiers: CD, DP, IE
2012 | Threatened – Nationally Vulnerable | Qualifiers: CD, IE
2009 | Threatened – Nationally Vulnerable | Qualifiers: IE, Inc, OL
2004 | Threatened – Nationally Critical
Referencing and citations
References and further reading
de Lange, P.J.; Cameron, E.K.; Murray, B.G. 1999: Alectryon excelsus subsp. grandis (Sapindaceae): a new combination for an uncommon small tree endemic to the Three Kings Islands, New Zealand. New Zealand Journal of Botany 37: 7-16.
Attribution
Description based on de Lange et al. (1999).
NZPCN Fact Sheet citation
Please cite as: de Lange, P.J. (Year at time of access): Alectryon excelsus subsp. grandis Fact Sheet (content continuously updated). New Zealand Plant Conservation Network. https://www.nzpcn.org.nz/flora/species/alectryon-excelsus-subsp-grandis/ (Date website was queried)