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  4. Parablechnum procerum

Parablechnum procerum

Tararua Range.<br>Photographer: Jeremy R. Rolfe, Date taken: 20/11/2011, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Tongariro.<br>Photographer: John Smith-Dodsworth, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Tongariro.<br>Photographer: John Smith-Dodsworth, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by-nc/4.0'>CC BY-NC</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Prothalli. In cultivation at Otari Wilton’s Bush, Wellington.<br>Photographer: Jeremy R. Rolfe, Date taken: 30/06/2011, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Prothalli. In cultivation at Otari Wilton’s Bush, Wellington.<br>Photographer: Jeremy R. Rolfe, Date taken: 30/06/2011, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
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Fertile frond amongst sterile fronds. Stokes Valley, Lower Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 08/01/2017, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Stokes Valley, Lower Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 08/01/2017, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Scales on rachis and abaxial surface of pinna. Stokes Valley, Lower Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 08/01/2017, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
Scales on fertile frond. Stokes Valley, Lower Hutt.<br>Photographer: Jeremy R. Rolfe, Date taken: 08/01/2017, Licence: <a target='_blank' href='https://creativecommons.org/licenses/by/4.0'>CC BY</a>. <a class='member-message' href='/nzpcn/why-join-nzpcn/' target='_blank'>Members can view a larger version of this image.</a>
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Common names

small kiokio

Biostatus

Native – Endemic taxon

Category

Vascular

Structural class

Ferns

Flower colours

No flowers

Detailed description

Rhizome short-creeping; scales to 16 × 2 mm wide, lanceolate, acute or acuminate, brown some bicolorous, entire. Fronds dimorphic, erect, 0.08–1.0 m long, 45–220 mm wide, widest mid frond; fertile fronds longer and more erect than sterile fronds. Stipes 20–600 mm (stipes up to 180 mm for plants growing in tussock communities, 150–250 for plants growing in open, shaded forest, and up to 0.6 m where plants are competing with a dense ground layer in the forest); stipes of sterile fronds shorter than stipes of fertile fronds, 1–4 mm diameter, stramineous to dark brown, becoming darker reddish brown towards the base; stipes scaly when young, often glabrous at maturity; scales 2–10 mm long, 0.5–1.5 mm wide, mostly lanceolate, brown, reddish brown, or somewhat bicolorous with paler margins, entire. Lamina ovate or lanceolate, dark olive green at maturity, 1-pinnate, 2–15 pairs of pinnae (plants suboptimal sites occasionally have as few as 1 terminal and 2 lateral, membranous pinnae), fertile laminae with more pinnae than sterile laminae. Rachis and costae dark, reddish, or pale brown (usually pale for specimens from open tussock communities), often blotchy; with sparse to numerous scales (fertile lamina with a more scaly rachis) and some small irregular hairs; scales variable in size, shape, and colour, 3.0–10.0 × 0.5–1.0 mm, linear to lanceolate, attenuate, more or less entire; conspicuous abaxial costal scales 2.0–3.5 mm long, c. 1 mm wide, of small narrow linear cells, lanceolate, attenuate, shiny dark reddish brown, concolorous or sometimes slightly bicolorous with paler margins (but lacking a “black-spot”), more or less entire. Sterile pinnae 25–150 × 10–35 mm, oblong-lanceolate, apices obtuse to acute, rounded or truncate at rachis; shortly petiolate at base of lamina, basiscopically adnate towards apex; coriaceous in robust specimens to almost membranous in small plants; margins toothed and sometimes crenate; veins simple or once-furcate; small toothed scales often extending on to lower surface of pinnae; basal pinnae as long or slightly 10–35 mm, oblong-lanceolate, apices obtuse to acute, rounded or truncate at rachis; shortly petiolate at base of lamina, basiscopically adnate towards apex; coriaceous in robust specimens to almost membranous in small plants; margins toothed and sometimes crenate; veins simple or once-furcate; small toothed scales often extending on to lower surface of pinnae; basal pinnae as long or slightly shorter than middle pinnae, rarely less than half their length, more obtuse, often reflexed, auricles and auriculate pinnae bases absent; terminal pinna usually longer and acutely pointed. Fertile pinnae 20–75 × 2.0–4.5 mm, linear, shortly petiolate at base of lamina, becoming basiscopically adnate towards apex; basal pinnae usually not reduced; sori covering under surface except at apices; indusium brown, laciniate or entire. Spores 56–70 × 39–52 µm.

Similar taxa

Parablechnum procerum is most similar to P. montanum, with which it sometimes hybridises. From P. montanum, P. procerum is distinguished by the usually fewer, blunter pinnae, basal pinnae which are not reduced. The pinnae of Parablechnum montanum are usually attenuate and falcate, and “black-spot” scales are present.

Distribution

Endemic. New Zealand: North Island (from about Mangamuka Forest south), South Island, Stewart Island/Rakiura, Chatham Islands and Sub-antarctic Islands. Becoming more common heading south and reaching sea level in the more southerly part of its range.

Habitat

Coastal to subalpine (montane to subalpine in northern part of range). Common in mixed forest, subalpine scrub and tussock grassland.

Current conservation status

The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2022-2023 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website. This report includes replaces all previous NZTCS lists for vascular plants. Previous assessments can be found here.

  • Conservation status of vascular plants in Aotearoa New Zealand, 2023. 2024. Peter J. de Lange, Jane Gosden, Shannel P. Courtney, Alexander J. Fergus, John W. Barkla, Sarah M. Beadel, Paul D. Champion, Rowan Hindmarsh-Walls, Troy Makan and Pascale Michel Department of Conservation. Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.

2023 | Not Threatened

Jump to previous conservation statuses

Detailed taxonomy

Genus

Parablechnum

Family

Blechnaceae

Authority

Parablechnum procerum (G.Forst.) C.Presl

Synonyms

Blechnum minus sensu Allan (1961); Stegania procera (G.Forst.) R.Br.; Onoclea procera (G.Forst.) Spreng.; Osmunda procera G.Forst.; Lomaria duplicata Potts; Lomaria latifolia Colenso; Lomaria procera (G.Forst.) Spreng.; Asplenium procerum (G.Forst.) Bernh.; Blechnopteris procera (G.Forst.) Trevis; Blechnum minus sensu Allan; Blechnum procerum (G.Forst.) Sw.

Taxonomic notes

Perrie et al. (2014) advocated for a broadened circumscription of Blechnaceae whereby a number of genera traditionally recognized as distinct from Blechnum were merged within it. However, this view has not met with universal acceptance (see Gasper et al. 2016) and does not seem to be followed worldwide (PPG 2016). From a New Zealand perspective the decision to merge Doodia in Blechnum, and rejection of Diploblechnum has not been universally accepted either e.g., Wilcox & Warden (2017), and as such it is considered appropriate to follow world opinion and accept the taxonomy of Gasper et al. (2016) and recommendations of the PPG (2016). See also the comments by Pyner (2017).

Endemic taxon

Yes

Endemic genus

No

Endemic family

No

Ecology

Flowering

Not applicable—spore producing

Fruiting

Not applicable—spore producing

Propagation technique

Easily grown from fresh spores and whole plants. transplants well and flourishes in most conditions but does best in a shaded site, planted in a fertile, permanently moist soil. Dislikes drought.

Wetland plant indicator status rating

Information derived from the revised national wetland plant list prepared to assist councils in delineating and monitoring wetlands (Clarkson et al., 2021 Manaaki Whenua – Landcare Research Contract Report LC3975 for Hawke’s Bay Regional Council). The national plant list categorises plants by the extent to which they are found in wetlands and not ‘drylands’. The indicator status ratings are OBL (obligate wetland), FACW (facultative wetland), FAC (facultative), FACU (facultative upland), and UPL (obligate upland). If you have suggestions for the Wetland Indicator Status Rating, please contact: [Enable JavaScript to view protected content]

FACU: Facultative Upland

Occasionally is a hydrophyte but usually occurs in uplands (non-wetlands).

Other information

Plant of the Month

This plant has been featured as a Plant of the Month – see Trilepidea: NZPCN newsletter for September 2017 for the full story.

Etymology

procerum: Long; from the Latin procerus

NVS code

The National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.

BLEPRO

Chromosome number

2n = 112

Previous conservation statuses

The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2022-2023 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website. This report includes replaces all previous NZTCS lists for vascular plants. Previous assessments can be found here.

  • Conservation status of vascular plants in Aotearoa New Zealand, 2023. 2024. Peter J. de Lange, Jane Gosden, Shannel P. Courtney, Alexander J. Fergus, John W. Barkla, Sarah M. Beadel, Paul D. Champion, Rowan Hindmarsh-Walls, Troy Makan and Pascale Michel Department of Conservation. Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.

2017 | Not Threatened

2012 | Not Threatened

2009 | Not Threatened

2004 | Not Threatened

Jump to current conservation status

Regional conservation statuses

Otago: 2024 | Regionally Not Threatened

The regional threat classification system leverages off the national assessments in the NZTCS, providing information relevant for the regional context. Otago conservation status information is sourced from the “Regional conservation status of indigenous vascular plants in Otago” Jarvie S et al. (2024) report.

Auckland: 2025 | At Risk – Regionally Naturally Uncommon | Qualifiers: DPR, DPS, DPT, RR

The regional threat classification system leverages off the national assessments in the NZTCS, providing information relevant for the regional context. Auckland conservation status information is sourced from the “Conservation status of vascular plant species in Tāmaki Makaurau / Auckland” Simpkins E et al. (2025) report.

Referencing and citations

References and further reading

Chambers TC, Farrant PA. 1998. The Blechnum procerum (“capense”) (Blechnaceae) complex in New Zealand. New Zealand Journal of Botany 36(1): 1–19. https://doi.org/10.1080/0028825X.1998.9512544.

Gasper AL, de Oliveira Dittrich VA, Smith AR, Salino A. 2016. A classification for Blechnaceae (Polypodiales: Polypodiopsida): New genera, resurrected names, and combinations. Phytotaxa 275: 191–227. https://doi.org/10.11646/phytotaxa.275.3.1.

Perrie LR, Wilson RK, Shepherd LD, Ohlsen DJ, Batty EL, Brownsey PJ, Bayly MJ. 2014. Molecular phylogenetics and generic taxonomy of Blechnaceae ferns. Taxon 63(4): 745–758. https://doi.org/10.12705/634.13.

PPG 1: The Pteridophyte Phylogeny Group 2016. A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution 54: 563–603. https://doi.org/10.1111/jse.12229.

Pyner T. 2017. A new classification of Blechnum. British Pteridological Society. https://ebps.org.uk/new-classification-blechnum/. Accessed [INSERT DATE ACCESSED].

Wilcox M, Warden J. 2017. Botany of Hillsborough coast bush reserves, Manukau Harbour, Auckland. Auckland Botanical Society Journal 72: 32–46.

Attribution

Fact sheet prepared for NZPCN by P.J. de Lange (7 March 2012). Description adapted Chambers & Farrant (1998)

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