Parablechnum procerum
Common names
small kiokio
Biostatus
Native – Endemic taxon
Category
Vascular
Structural class
Ferns
Flower colours
No flowers
Detailed description
Rhizome short-creeping; scales to 16 × 2 mm wide, lanceolate, acute or acuminate, brown some bicolorous, entire. Fronds dimorphic, erect, 0.08–1.0 m long, 45–220 mm wide, widest mid frond; fertile fronds longer and more erect than sterile fronds. Stipes 20–600 mm (stipes up to 180 mm for plants growing in tussock communities, 150–250 for plants growing in open, shaded forest, and up to 0.6 m where plants are competing with a dense ground layer in the forest); stipes of sterile fronds shorter than stipes of fertile fronds, 1–4 mm diameter, stramineous to dark brown, becoming darker reddish brown towards the base; stipes scaly when young, often glabrous at maturity; scales 2–10 mm long, 0.5–1.5 mm wide, mostly lanceolate, brown, reddish brown, or somewhat bicolorous with paler margins, entire. Lamina ovate or lanceolate, dark olive green at maturity, 1-pinnate, 2–15 pairs of pinnae (plants suboptimal sites occasionally have as few as 1 terminal and 2 lateral, membranous pinnae), fertile laminae with more pinnae than sterile laminae. Rachis and costae dark, reddish, or pale brown (usually pale for specimens from open tussock communities), often blotchy; with sparse to numerous scales (fertile lamina with a more scaly rachis) and some small irregular hairs; scales variable in size, shape, and colour, 3.0–10.0 × 0.5–1.0 mm, linear to lanceolate, attenuate, more or less entire; conspicuous abaxial costal scales 2.0–3.5 mm long, c. 1 mm wide, of small narrow linear cells, lanceolate, attenuate, shiny dark reddish brown, concolorous or sometimes slightly bicolorous with paler margins (but lacking a “black-spot”), more or less entire. Sterile pinnae 25–150 × 10–35 mm, oblong-lanceolate, apices obtuse to acute, rounded or truncate at rachis; shortly petiolate at base of lamina, basiscopically adnate towards apex; coriaceous in robust specimens to almost membranous in small plants; margins toothed and sometimes crenate; veins simple or once-furcate; small toothed scales often extending on to lower surface of pinnae; basal pinnae as long or slightly 10–35 mm, oblong-lanceolate, apices obtuse to acute, rounded or truncate at rachis; shortly petiolate at base of lamina, basiscopically adnate towards apex; coriaceous in robust specimens to almost membranous in small plants; margins toothed and sometimes crenate; veins simple or once-furcate; small toothed scales often extending on to lower surface of pinnae; basal pinnae as long or slightly shorter than middle pinnae, rarely less than half their length, more obtuse, often reflexed, auricles and auriculate pinnae bases absent; terminal pinna usually longer and acutely pointed. Fertile pinnae 20–75 × 2.0–4.5 mm, linear, shortly petiolate at base of lamina, becoming basiscopically adnate towards apex; basal pinnae usually not reduced; sori covering under surface except at apices; indusium brown, laciniate or entire. Spores 56–70 × 39–52 µm.
Similar taxa
Parablechnum procerum is most similar to P. montanum, with which it sometimes hybridises. From P. montanum, P. procerum is distinguished by the usually fewer, blunter pinnae, basal pinnae which are not reduced. The pinnae of Parablechnum montanum are usually attenuate and falcate, and “black-spot” scales are present.
Distribution
Endemic. New Zealand: North Island (from about Mangamuka Forest south), South Island, Stewart Island/Rakiura, Chatham Islands and Sub-antarctic Islands. Becoming more common heading south and reaching sea level in the more southerly part of its range.
Habitat
Coastal to subalpine (montane to subalpine in northern part of range). Common in mixed forest, subalpine scrub and tussock grassland.
Current conservation status
The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2022-2023 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website. This report includes replaces all previous NZTCS lists for vascular plants. Previous assessments can be found here.
Please note, threat classifications are often suggested by authors when publications fall between NZTCS assessment periods – these interim threat classification statuses has not been assessed by the NZTCS panel.
- Conservation status of vascular plants in Aotearoa New Zealand, 2023. 2024. Peter J. de Lange, Jane Gosden, Shannel P. Courtney, Alexander J. Fergus, John W. Barkla, Sarah M. Beadel, Paul D. Champion, Rowan Hindmarsh-Walls, Troy Makan and Pascale Michel Department of Conservation. Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.
2023 | Not Threatened
Detailed taxonomy
Family
Blechnaceae
Synonyms
Blechnum minus sensu Allan (1961); Stegania procera (G.Forst.) R.Br.; Onoclea procera (G.Forst.) Spreng.; Osmunda procera G.Forst.; Lomaria duplicata Potts; Lomaria latifolia Colenso; Lomaria procera (G.Forst.) Spreng.; Asplenium procerum (G.Forst.) Bernh.; Blechnopteris procera (G.Forst.) Trevis; Blechnum minus sensu Allan; Blechnum procerum (G.Forst.) Sw.
Taxonomic notes
Perrie et al. (2014) advocated for a broadened circumscription of Blechnaceae whereby a number of genera traditionally recognized as distinct from Blechnum were merged within it. However, this view has not met with universal acceptance (see Gasper et al. 2016) and does not seem to be followed worldwide (PPG 2016). From a New Zealand perspective the decision to merge Doodia in Blechnum, and rejection of Diploblechnum has not been universally accepted either e.g., Wilcox & Warden (2017), and as such it is considered appropriate to follow world opinion and accept the taxonomy of Gasper et al. (2016) and recommendations of the PPG (2016). See also the comments by Pyner (2017).
Endemic taxon
Yes
Endemic genus
No
Endemic family
No
Ecology
Flowering
Not applicable—spore producing
Fruiting
Not applicable—spore producing
Propagation technique
Easily grown from fresh spores and whole plants. transplants well and flourishes in most conditions but does best in a shaded site, planted in a fertile, permanently moist soil. Dislikes drought.
Wetland plant indicator status rating
Information derived from the revised national wetland plant list prepared to assist councils in delineating and monitoring wetlands (Clarkson et al., 2021 Manaaki Whenua – Landcare Research Contract Report LC3975 for Hawke’s Bay Regional Council). The national plant list categorises plants by the extent to which they are found in wetlands and not ‘drylands’. The indicator status ratings are OBL (obligate wetland), FACW (facultative wetland), FAC (facultative), FACU (facultative upland), and UPL (obligate upland). If you have suggestions for the Wetland Indicator Status Rating, please contact: [Enable JavaScript to view protected content]
FACU: Facultative Upland
Occasionally is a hydrophyte but usually occurs in uplands (non-wetlands).
Other information
Etymology
procerum: Long; from the Latin procerus
NVS code
The National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.
BLEPRO
Chromosome number
2n = 112
Previous conservation statuses
2017 | Not Threatened
2012 | Not Threatened
2009 | Not Threatened
2004 | Not Threatened
Referencing and citations
References and further reading
Chambers TC, Farrant PA. 1998. The Blechnum procerum (“capense”) (Blechnaceae) complex in New Zealand. New Zealand Journal of Botany 36(1): 1–19. https://doi.org/10.1080/0028825X.1998.9512544.
Gasper AL, de Oliveira Dittrich VA, Smith AR, Salino A. 2016. A classification for Blechnaceae (Polypodiales: Polypodiopsida): New genera, resurrected names, and combinations. Phytotaxa 275: 191–227. https://doi.org/10.11646/phytotaxa.275.3.1.
Perrie LR, Wilson RK, Shepherd LD, Ohlsen DJ, Batty EL, Brownsey PJ, Bayly MJ. 2014. Molecular phylogenetics and generic taxonomy of Blechnaceae ferns. Taxon 63(4): 745–758. https://doi.org/10.12705/634.13.
PPG 1: The Pteridophyte Phylogeny Group 2016. A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution 54: 563–603. https://doi.org/10.1111/jse.12229.
Pyner T. 2017. A new classification of Blechnum. British Pteridological Society. https://ebps.org.uk/new-classification-blechnum/. Accessed [INSERT DATE ACCESSED].
Wilcox M, Warden J. 2017. Botany of Hillsborough coast bush reserves, Manukau Harbour, Auckland. Auckland Botanical Society Journal 72: 32–46.
Attribution
Fact sheet prepared for NZPCN by P.J. de Lange (7 March 2012). Description adapted Chambers & Farrant (1998)
NZPCN Fact Sheet citation
Please cite as: de Lange, P.J. (Year at time of access): Parablechnum procerum Fact Sheet (content continuously updated). New Zealand Plant Conservation Network. https://www.nzpcn.org.nz/flora/species/parablechnum-procerum/ (Date website was queried)