Leptinella atrata subsp. atrata

Detailed description

Fleshy, monoecious, perennial herb producing 1 or more summer green leaf tufts amongst mobile scree. Rhizomes ascending, at or close to rock surface when young, becoming deeply buried with age, up to 10 mm long, thick, very fleshy, pale or pinkish-red, sparsely short-hairy, glabrate; branches often in clusters of up to 4 diverging from a flowering node and the nodes immediately behind, most of these dying in the first season, leaves usually crowded around rhizome apex, sometimes up to 25 mm apart. Roots numerous, fleshy, very long, extensively branching, thick up to 2 mm diameter. leaves 2-pinnatifid, 20-80 x 5-13 mm; blade 15-60 mm long, obovate, coriaceous and fleshy, grey-green tinged with red, especially on the primary and secondary axes, sparsely pilose hairy, midrib not raised on ventral surface; pinnae 8-15 pairs, cut to rhachis, proximal pinnae distant, distal ones overlapping, broadly elliptic and divided, reducing to distant oblong simple proximal pinnae; secondary pinnae 0-9 per pinna, equally on distal and proximal sides, cut to midrib of pinna, oblong or obovate, with rounded apices, upturned, occasionally with 1-2 small lobes on either side. Peduncles scarcely longer than leaves, 30-120 mm, stout, fleshy, pilose hairy with 4-10 evenly spaced bracts; lowermost bracts deeply divided, scarcely smaller than leaves, reducing evenly to simple, oblong or shallowly divided uppermost bracts. Capitula 5-13 mm diameter, surface convex; involucre hemispherical; involucral bracts unequally 2-3-seriate, overtopped by mature florets, grey-green tinged pink or red, more or less pilose hairy, outer bracts exceeding florets, thick, pinnatifid with up to 6 pairs of oblong lobes on sides, gradually changing to thinner, simple obovate inner involucral bracts whose margins are finely scarious; receptacle convex; pistillate florets 100-240 in several rows, 3.25-3.75 mm long, straight, very dark red to almost black; corolla 5 times as long as wide, teeth conspicuous, equal and diverging; staminate florets equal in number. Stigmas of pistillate and staminate florets not exserted far beyond corolla mouth, retracting into corolla tube at anthesis. Cypsela 1.8-2.8 x 0.8-1 mm, slightly compressed, pale brown or dark brown, deeply wrinked when mature