Veronica pinguifolia
Common name
hebe
Synonyms
Hebe pinguifolia (Hook.f.) Cockayne et Allan
Family
Plantaginaceae
Flora category
Vascular – Native
Endemic taxon
Yes
Endemic genus
No
Endemic family
No
Structural class
Trees & Shrubs - Dicotyledons
NVS code
The National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.
HEBPIN
Chromosome number
2n = 80
Current conservation status
The threat classification status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2017 using the New Zealand Threat Classification System (NZTCS) – more information about this can be found on the NZTCS website This report includes a statistical summary and brief notes on changes since 2012 and replaces all previous NZTCS lists for vascular plants. Authors: By Peter J. de Lange, Jeremy R. Rolfe, John W. Barkla, Shannel P. Courtney, Paul D. Champion, Leon R. Perrie, Sarah M. Beadel, Kerry A. Ford, Ilse Breitwieser, Ines Schönberger, Rowan Hindmarsh-Walls, Peter B. Heenan and Kate Ladley. Please note, threat classifications are often suggested by authors when publications fall between NZTCS assessment periods – a suggested threat classification status has not been assessed by the NZTCS panel.
Source: NZTCS and licensed by DOC for reuse under the Creative Commons Attribution 4.0 International licence.
2017 | Not Threatened
Previous conservation statuses
2012 | Not Threatened
2009 | Not Threatened
2004 | Not Threatened
Brief description
Low growing bushy shrub beating pairs of thick blue-green strongly dished and often red-edged oval leaves on an erect reddish stem. Leaves 7-16mm long, variable in width. Leaf bud without gap at base. Flowers white, in a spike to 2cm long towards the tip of twigs.
Distribution
South Island - Mountains east of the Main Divide, from the Bryant Range to the Kakanui Mountains
Habitat
Open alpine areas, on rocks and debris slopes, sometimes in grassland.
Features
Spreading low shrub (openly branched, or compact) to 0.4 (-0.8) m tall. Branches decumbent or spreading or erect, old stems dark brown or grey; branchlets green (tinged maroon, especially at nodes) or red-brown, puberulent, hairs bifarious; internodes (0.5-) 1-7 (-10) mm; leaf decurrencies evident. Leaf bud distinct; sinus absent (usually), or small and acute to rounded. Leaves erect or erecto-patent; lamina lanceolate (often broadly) to ovate or obovate, fleshy, concave, (3-) 7-16 (-22) x (2-) 4-9 (-12) mm; apex rounded or sometimes subacute; midrib very slightly thickened below; margin usually minutely papillate and rarely glandular-ciliate (toward leaf base), often tinged red; upper surface glaucous (usually) or glaucescent, with many stomata, glabrous; lower surface glaucous (usually) or glaucescent. Inflorescences with ( 4-) 12-22 flowers, lateral, unbranched, 1-2.8 (-3.4) cm, about equal to or longer than subtending leaves; peduncle 0.3-1.5 (-2.1) cm; rachis 0.3-1 (-1.5) cm. Bracts opposite and decussate (or apparently so) or lowermost pair opposite, then subopposite or alternate above, ovate (often narrowly) or deltoid, subacute. Flowers hermaphrodite or female (on different plants). Pedicels absent or if evident then always shorter than bracts, 0-0.8 mm. Calyx (2-) 2.7- 3.2 (-4) mm; lobes elliptic or oblong or ovate, subacute to obtuse. Corolla tube glabrous; tube of hermaphrodite flowers 2-3 x approximately 1-1.5 mm, cylindric or narrowly funnelform, approximately equalling calyx; tube of female flowers 2-2.5 x approximately 1.5-1.8 mm, cylindric or narrowly funnelform, approximately equalling calyx; lobes white at anthesis, ovate or lanceolate or elliptic, obtuse, suberect to recurved, longer than corolla tube. Stamen filaments 4.5-5 mm; anthers magenta, 2.1-2.3 mm; sterile anthers of female flowers magenta or buff, 1.4-1.6 mm. Ovary ovoid or globose, hairy, 0.5-1.1 mm, apex (in septum view) obtuse or slightly didymous; ovules approximately 8-13 per locule, in 1 layer (but sometimes a few more or less overlapping); style (4-) 5-7.5 mm, hairy. Capsules obtuse or truncate, 3-4.5 x 2.5-3.2 mm, usually hairy, loculicidal split extending ¼-way to base. Seeds flattened, ellipsoid to oblong, more or less smooth, brown (sometimes pale), 0.9-1.7 x 0.6-1.1 mm, micropylar rim 0.3-0.6 mm.
Similar taxa
A variable species, distinguished from most others by the combination of the shape and size of the glaucous leaves, glabrous leaf margins, sessile flowers, and the length of bracts relative to calyces. The limits of the species are not well defined, and differences from V. buchananii (see Taxonomic Notes below) are problematic. It has sometimes been confused with V. baylyi (see Taxonomic Notes below), and specimens are sometimes misidentified as V. decumbens and vice versa (see notes under that species).
No single character has been found to distinguish V. pinguifolia and V. buchananii consistently, and they are generally distinguished here on combinations of characters. V. pinguifolia plants are often taller (although sprawling, they do not tend to form dense mats) and usually have more distinct leaf buds, these not closely surrounded by recently diverged leaf pairs (except in some Marlborough specimens). They mostly have larger leaves (although shape is variable) that are not keeled when fresh (although they may appear so when dry, as the fleshy lamina shrinks away from the midrib). They may have more slender, less corky stems, and bracts and calyces that are usually shortly ciliolate with glandular hairs (but sometimes long-ciliate with eglandular hairs). In contrast, V. buchananii tends to be more mat-forming (except for “var. exigua-like” plants) and lower growing, with leaf buds closely surrounded by recently diverged leaves. It often has smaller leaves (although shape is variable) that are more keeled. It also often has thicker, more corky stems, and has calyces and bracts that often have longer cilia.
Flowering
December-February (-April)
Flower colours
Cream, White
Fruiting
January-April
Life cycle
Seeds are wind dispersed (Thorsen et al., 2009).
Etymology
veronica: Named after Saint Veronica, who gave Jesus her veil to wipe his brow as he carried the cross through Jerusalem, perhaps because the common name of this plant is ‘speedwell’. The name Veronica is often believed to derive from the Latin vera ‘truth’ and iconica ‘image’, but it is actually derived from the Macedonian name Berenice which means ‘bearer of victory’.
pinguifolia: From the Latin pinguis ‘fat, plump’ and folium ‘leaf’, in reference to the thickness of the leaves
Taxonomic notes
Some specimens of V. pinguifolia / V. buchananii have not been identified with certainty, and the distribution maps for both species are based only on specimens about whose identities we are reasonably confident. As defined here, there is some geographic overlap between the two species. Further investigation of their variation, relationships and circumscriptions would be worthwhile.
Included here in V. pinguifolia are specimens from the north of the species’ range (e.g. Mount Starveall, Travers Range, St Arnaud Range, Hodder Valley, Black Birch Range) that sometimes have a small but distinct sinus in the leaf bud (a feature seen only rarely on plants from other areas). In this respect, these specimens resemble V. baylyi, a name that has sometimes been applied to them. They can be distinguished from that species by usually red-edged leaves that are paler green (under a glaucous bloom), larger, strictly opposite bracts, and blunt and usually hairy capsules. Some of these northern specimens (particularly those from Black Birch Range) are quite small-leaved and, in this respect, may also resemble V. buchananii. However, given their geographic distance from that species, the resemblance is probably coincidental, and a close relationship does not seem likely.
Both diploid and tetraploid chromosome numbers are recorded in V. pinguifolia, but chromosome variation has not been correlated with variation in morphology. Some vouchers for chromosome counts (diploid from Mount Peel, Canterbury; tetraploid from Mount Somers and Mount Winterslow) have been identified here as V. cf. pinguifolia. These specimens are cultivated and sterile and cannot be identified with certainty. V. pinguifolia does, on the basis of other specimens, occur on Mount Peel and Mount Somers but is not otherwise known from Mount Winterslow.
Attribution
Description adapted by M. Ward from Bayly & Kellow (2006).
References and further reading
Bayly, M.J., Kellow, A.V. 2006. An illustrated guide to New Zealand Hebes. Wellington, N.Z.: Te Papa press pg. 136-138.
Thorsen, M. J.; Dickinson, K. J. M.; Seddon, P. J. 2009. Seed dispersal systems in the New Zealand flora. Perspectives in Plant Ecology, Evolution and Systematics 11: 285-309
NZPCN Fact Sheet citation
Please cite as: Ward, M.D. (Year at time of access): Veronica pinguifolia Fact Sheet (content continuously updated). New Zealand Plant Conservation Network. https://www.nzpcn.org.nz/flora/species/veronica-pinguifolia/ (Date website was queried)